1948, B.A. in Biology, Goshen College, Goshen, Indiana.

1953, Ph.D. in Zoology, Eigenmann Fellow, Indiana University, Bloomington, Indiana.

1955-57, Post-doctoral studies, Duke University, Durham, North Carolina.

1957-61, Associate Professor of Biology, Bethany College, Bethany, West Virginia.

1961-64, Professor, Eastern Mennonite College, Harrisonburg, Virginia.

1964-86, Research Professor in Zoology, Goshen College, Goshen, Indiana.

1986-Present, Research Professor Emeritus, Goshen College, Goshen, Indiana.